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那么我們熟悉的中相手蟹/仿相手蟹去哪了呢？

中相手蟹Sesarmops于Serène 和 Soh 在1970年對于東太平洋和印度洋的相手蟹類建立的一個屬，其中包括了六個物種，隨后這些物種都被慢慢的歸于其他的屬了，比如S. sinensis (H. Milne Edwards, 1853) 和?S. intermedius?(De Haan, 1835)以及被轉(zhuǎn)為中華東方相手Orisarma sinense (H. Milne Edwards, 1853) 和中型東方相手Orisarma intermedium (De Haan, 1835) 。

定義了 Sesarmops如下：甲殼長或長于寬度在外眶齒處測量，額緣有一個很深的中凹，后額葉是突出，外眶齒明顯分離與側(cè)緣的其余部分相距明顯的裂縫，與胃和心臟區(qū)域明確界定。

一般的該屬成員大抵是這個形象的。

中相手蟹屬Sesarmops Serène?&?Soh, 1970

屬內(nèi)一共8種。

模式種：印痕相手蟹Sesarma impressus H. Milne Edwards, 1837=印痕中相手蟹Sesarmops impressus (H. Milne Edwards, 1837)

紫黑中相手蟹Sesarmops atrorubens (Hess, 1865)

Sesarma atrorubens Hess, 1865: 149, pl. 6 fig. 12 (Sydney); Haswell? 1882: 108 (list); De Man 1887: 653, 676, 678 (part) (Sydney); De Man 1890: 95 (Fiji); Ortmann 1894: 724 (Fiji).

Sesarma (Sesarma) atrorubens – Tesch 1917: 131 (part) (Fiji).

Sesarmops impressum – Serène and Soh 1970: 401, 406 (list). (not Sesarma impressa H. Milne Edwards, 1837).

Sesarma (Sesarmops) impressum – McLay and Ryan 1990: 108, figs. 1A–G (Fiji). (not Sesarma impressa H. Milne Edwards, 1837).

Sesarmops impressus – Ng et al. 2008: 223 (part; list). (not Sesarma?

impressa H. Milne Edwards, 1837).

幾乎僅見于斐濟(jì)。

種加詞意為黑紅色的，不過活體顏色顯然更偏紫色一些。

本種分布相對狹窄，但是體色十分美麗。

帝王中相手蟹Sesarmops imperator Ng, Li & Shih, 2020

?Sesarma atrorubens – De Man 1887: 653, 676, 678 (part) (Timor, ?Ambon and Xulla Besi [Moluccas], Sanghir [northern ?Sulawesi]).?

Sesarma impressa – De Man 1892: 330 (Timor); Bürger 1893: 620, ?pl. 21 figs. 4, 5 (Philippines; Palau Islands); Schenkel 1902: 546 ?(Sulawesi). (not Sesarma impressa H. Milne Edwards, 1837)?

Sesarma frontalis – De Man 1892: 334, pl. 19 fig. 13 (Flores). (not ?Sesarma frontale A. Milne-Edwards, 1869).?

?Sesarma (Sesarma) impressa – Nobili 1900: 507 (Timor).?

Sesarma (Sesarma) impressa – De Man 1902: 527 (Ternate, Batjan, ?Halmahera); Tesch 1917: 158 (Halmahera, Sulawesi, Nias, ?Papua New Guinea); Balss 1922: 155 (southern Taiwan); Sakai ?1939: 685, pl. 110 fig. 1 (Danshuei, Taiwan); Horikawa 1940: ?30 (Taiwan; list); Sakai 1940: 32 (Taiwan; list); Lin 1949: 30 ?(southern Taiwan; no new specimens); Shen and Dai 1964: 136, ?1 unnumbered fig. (Hainan, China). (not Sesarma impressa H. ?Milne Edwards, 1837).?

?Sesarma (Sesarma) atrorubens – Tesch 1917: 131 (part) (Timor, ?Ambon and Soela Besi [Moluccas], Sanghir [northern ?Sulawesi]).?

Sesarma (Parasesarma) impressa – Miyake 1939: 226, 244 (Palau). ?(not Sesarma impressa H. Milne Edwards, 1837).?

Sesarma (Sesarmops) impressum – Dai et al. 1986: 493, pl. 69(7), ?text fig. 178(1–3) (Taiwan); Dai and Yang 1991: 540, pl. 69(7), ?text fig. 178(1–3) (Taiwan). (not Sesarma impressa H. Milne ?Edwards, 1837).?

Sesarmops impressum – Serène and Soh 1970: 401, 406 (part), pl. ?7 figs. A, B (no locality data); Sakai 1976: 660, text fig. 360 ?(Taiwan); Wang and Liu 1996: 123, figs. 161–162 (Taiwan); Liu ?1999: 88 (Taiwan); Cai and Ng 2001: 691, fig. 19 (Halmahera, ?Indonesia); Lee 2001: 134, 2 unnumbered figs. (Taiwan); Ng et ?al. 2001: 43 (Taiwan); C.-H. Jeng and M.-F. Wang 2002: 85, 2 ?unnumbered figs. (Taiwan); Ho 2003: 9, 34, 2 unnumbered figs. ?(Taiwan). Liu 2009: 57, fig. 18 (Taiwan); Li and Chiu 2013: ?56, 3 unnumbered figs. (Taiwan); Li and Chiu 2019a: 89; Li ?and Chiu 2019b: 45 (Taiwan); Ng et al. 2017: 106; Shao et al. ?2007: 207, 1 unnumbered fig. (not Sesarma impressa H. Milne ?Edwards, 1837).?

Parasesarma pictum – Y.-H. Chen 2001: 222, 1 unnumbered fig. ?[lower]. (not Grapsus (Pachysoma) pictus De Haan, 1835)?

Sesarma impressum – W.-J. Chen and Lo 2014: 144, 2 unnumbered ?figs. (Taiwan). (not Sesarma impressa H. Milne Edwards, 1837)?

Sesarmops impressus – Fukui et al. 1989: 230 (Taiwan); Yu et al.1996: 15, figs. 75–76 (Taiwan); Jeng 1997: 18 (Taiwan); Jeng 1998: lower image on p. 81, upper image on p. 82 (Taiwan); Ng et al. 2008: 223 (part); C.-H. Jeng et al. 2010: 72, 5 unnumbered figs. (Taiwan); Li et al. 2010: 6 (Taiwan); Liu and Wang 2010: 61, 3 unnumbered figs. (Taiwan); Maenosono and Naruse 2016: 7 (Ryukyus, Japan; Lanyu, Taiwan); Maenosono and Saeki 2016: 9 (Ryukyus, Japan). (not Sesarma impressa H. Milne Edwards, 1837).

Sesarmops impressus 2 – Li et al. 2020: fig. 22 (Taiwan; Philippines; Sulawesi, Indonesia).

帝王是我迪亞波羅噠！

早期認(rèn)為是印痕，后來證實完全不是一個東西（顏色都差很多），通常頭胸甲看起來是紫褐色的，兩側(cè)和前部的邊緣金黃色，螯足紫紅色或淡紫色，密布白色的小顆粒。

種名由于紫色是歐洲（精羅落淚）皇室的代表。

該種最典型的特點是頭胸甲長明顯大于寬且前部稍窄，臺灣報道很多，喜歡生活在通海溪流的附近，經(jīng)常可以見到較大群，不過有趣的是相較于東方相手蟹屬，該種喜歡郁閉度更高環(huán)境，而且更喜歡淡水。

印痕中相手蟹Sesarmops impressus (H. Milne Edwards, 1837)

Sesarma impressa H. Milne Edwards, 1837: 74 (no locality); H. ?Milne Edwards 1853: 186 (no locality); De Man 1887: 653, 671 ?(Madagascar); Lenz 1905: 370 (Zanzibar).?

Sesarma frontale A. Milne-Edwards, 1869: 27 (Madagascar).?

Sesarma frontalis – De Man 1887: 649 (Madagascar).?

Sesarma nodulifera – Lenz 1910: 562 (Comoros).?

?Sesarma impressa – Lenz 1910: 561 (Madagascar).

Sesarma (Sesarma) impressum – Crosnier 1965: 63, figs. 82, 92, 104 ?(Madagascar, Comoros); Serène 1968: 106.?

Sesarma (Sesarma) impressa – Guinot 1967: 287 (list, Comoros).?

Sesarmops impressum – Serène and Soh 1970: 401, 406 (part), pl. 7 ?figs. A, B (no locality data).?

Sesarmops impressus – Keith et al. 2006: 36 (Comoros, Mayotte); ?Ng et al. 2008: 223 (list); Bouchard 2009: 6, 15 (Mayotte); ?Bouchard et al. 2013: 24, fig. 19 (Mayotte); Poupin et al. 2018: ?72 (Mayotte). Sesarmops impressus 1 – Li et al. 2020: fig. 22 (Mayotte; ?Madagascar).

非洲東海岸，似乎更喜歡島嶼上的濕潤環(huán)境，生活在溪流邊。成年雄性具有對比度極大的配色，螯足及頭胸甲的前部均為灰綠色至污白色，其余部分棕黑色或深黑色。

印度中相手蟹Sesarmops indicus Ng, Li & Shih, 2020

?Sesarma (Episesarma) frontalis – De Man 1895: 172 (Atjeh, ?northern Sumatra).?

?Sesarma (Sesarma) impressa – Nobili 1900: 507 (Mentawei Islands, ?west coast Sumatra).?

?Sesarma (Sesarma) frontalis – Nobili 1900: 509 (Engano, west coast ?Sumatra).

該種僅見于安達(dá)曼海，就是說馬爾代夫應(yīng)當(dāng)有分布。但是實際上好像沒有什么活體圖可以看···標(biāo)本都是1998年采集的。

棉蘭老中相手蟹Sesarmops mindanaoensis (Rathbun, 1914)

Sesarma (Sesarma) mindanaoense Rathbun, 1914: 75.

Sesarmops mindanaoensis – Serène and Soh 1970: 401, 406; Ng et al. 2008: 223; Liu 2013: 32, fig. 2.

目前報道于菲律賓和臺灣，這個種顏色相對灰暗，行蹤更隱秘。

延遲中相手蟹Sesarmops mora Li, Shih & Ng, 2020

Sesarma frontalis – De Man 1892: 334, pl. 19(13). (not Sesarma ?frontale A. Milne-Edwards, 1869 = Sesarma impressa H. Milne ?Edwards, 1837).

見于臺灣、菲律賓和印尼蘇拉威西，該種因為拖延了很久才發(fā)表而得名，可見作者怨念.jpg

螯足紫紅色，在身體后部向前延伸至中胃區(qū)兩側(cè)淺色，形成一個A字型條帶。

相似中相手蟹Sesarmops similis (Hess, 1865)

Sesarma similis Hess, 1865: 150 (“Sydney”); Haswell 1882: 108 (list).?

Sesarma impressa – Ortmann 1894: 723 (Samoa). (not Sesarma impressa H. Milne Edwards, 1837).

本種可能僅見于薩摩亞，看起來應(yīng)當(dāng)是喜歡太平洋海島上的環(huán)境。

韋氏中相手蟹Sesarmops weberi (De Man, 1892)

Sesarma weberi - de Man, 1892:338-341 Pls. XX-XXI（Fig.15）

Sesarmops weberi -?Ng ?et al. 2008: 223

Sesarmops weberi -?Li et al. 2020:10, 16

該種分布于弗洛勒斯島，具體情況不明，也沒有圖，De Man本人記載是淡水河的瀑布下面采的。

不過，該屬部分種類和針肢蟹屬具有非常近的演化關(guān)系，加之南部的種類生殖器形態(tài)還真的就和北部的不一樣，顯然之后可能會進(jìn)行調(diào)整。

未完待續(xù)

借物表

Ng, P. K., Li, J. J., & Shih, H. T. (2020). What is Sesarmops impressus (H. Milne Edwards, 1837)(Crustacea: Brachyura: Sesarmidae)?. Zoological Studies, 59.

Li, J. J., Shih, H. T., & Ng, P. K. (2020). The Taiwanese and Philippine species of the terrestrial crabs Bresedium Serène and Soh, 1970 and Sesarmops Serène and Soh, 1970 (Crustacea: Decapoda: Brachyura), with descriptions of two new species. Zoological Studies, 59.